Perennial herbs; stems 40-80 cm tall, slightly woody at base, nodes sometimes inflated, mostly pubescent; leaves narrowly or broadly ovate, glabrescent or pubescent, base oblique, cuneate, margin entire or coarsely dentate, sometimes with salient, unequal deltoid lobes, apex acuminate; pedicel glabrescent, puberulent or densely and persistently villous; calyx broadly campanulate, densely pubescent; corolla whitish yellow, with a greenish yellow or darker eye near the base, rotate or campanulate; stamens yellow attached near the base of corolla, anthers oblong, shorter than filaments; ovary 2-celled, many ovuled, style linear, stigma slightly 2-lobed; fruiting calyx orange or red, ovate, rounded, 10-ribbed, subleathery, invaginated at base, with glabrescent or persistent villous; berry shiny, orange-red, globose; seeds small, pale yellow, reniform.